diff --git a/.changeset/projection-staged-error-reader-link-4517.md b/.changeset/projection-staged-error-reader-link-4517.md
new file mode 100644
index 0000000000..5170e6c34b
--- /dev/null
+++ b/.changeset/projection-staged-error-reader-link-4517.md
@@ -0,0 +1,5 @@
+---
+"@solidjs/signals": patch
+---
+
+A reader of a derived store whose derive carries a staged error (D-41) now links to the derive, as a reader of a pending or errored one does, so it re-runs when the derive's source answers. An `<Errored>` mounted in the change that errored its content over a keyed `createProjection` showed its fallback for good; it now shows the answer.
diff --git a/packages/signals/src/store/store.ts b/packages/signals/src/store/store.ts
index d12476c0fd..9d318b674a 100644
--- a/packages/signals/src/store/store.ts
+++ b/packages/signals/src/store/store.ts
@@ -36,6 +36,7 @@ import {
   CONFIG_MANUAL_WRITE,
   CONFIG_OVERRIDE,
   CONFIG_PROMOTED,
+  CONFIG_STAGED_ERROR,
   CONFIG_VERDICT,
   EFFECT_RENDER,
   NOT_PENDING,
@@ -1844,7 +1845,8 @@ function pullFamily(target: StoreTarget): void {
   // reader of a memo would — linked, so it re-runs at the landing (the
   // next settled pass pulls without linking and the stale link trims), a
   // verdict reader registered, a tracked pass suspended.
-  if (fw._statusFlags & (STATUS_PENDING | STATUS_ERROR)) {
+  // A staged error (D-41) throws at the pull, before any leaf would link the reader.
+  if (fw._statusFlags & (STATUS_PENDING | STATUS_ERROR) || fw._config & CONFIG_STAGED_ERROR) {
     // A render effect outside the flight's own flush is the frame, not a
     // derivation: it keeps what it shows and learns of the landing from the
     // leaves the landing changes (unchanged leaves say nothing — it is not a
@@ -1858,7 +1860,7 @@ function pullFamily(target: StoreTarget): void {
     if (
       c !== null &&
       c._type === EFFECT_RENDER &&
-      !(fw._statusFlags & STATUS_ERROR) &&
+      fw._statusFlags & STATUS_PENDING &&
       fw._config & CONFIG_HELD &&
       flushTransaction !== txOf(fw) &&
       !linkedTo(c, fw)
diff --git a/packages/signals/tests/errored-born-staged-error-projection-4517.test.ts b/packages/signals/tests/errored-born-staged-error-projection-4517.test.ts
new file mode 100644
index 0000000000..1fb438ad01
--- /dev/null
+++ b/packages/signals/tests/errored-born-staged-error-projection-4517.test.ts
@@ -0,0 +1,86 @@
+import { expect, it } from "vitest";
+import {
+  createErrorBoundary,
+  createLoadingBoundary,
+  createMemo,
+  createProjection,
+  createRenderEffect,
+  createRoot,
+  createSignal,
+  flush,
+  mapArray,
+  untrack
+} from "../src/index.js";
+
+const tick = async () => {
+  for (let i = 0; i < 4; i++) {
+    await new Promise(r => setTimeout(r, 0));
+    flush();
+  }
+};
+
+type Row = { id: number; name: string };
+
+for (const keyed of [true, false])
+  it(`an answer replaces the error a boundary was born with (projection: ${keyed}) (#4517)`, async () => {
+    const asks: { key: string; res: (v: Row[]) => void }[] = [];
+    const ask = (key: string) => new Promise<Row[]>(res => asks.push({ key, res }));
+    const land = async (key: string, v: Row[]) => {
+      for (const a of asks.filter(a => a.key === key)) {
+        asks.splice(asks.indexOf(a), 1);
+        a.res(v);
+      }
+      await tick();
+    };
+    const [q, setQ] = createSignal("");
+    let screen: unknown;
+    const dispose = createRoot(d => {
+      const src = createMemo(() => {
+        const s = q();
+        if (s === "") return [] as Row[];
+        if (s.length < 2) throw new Error("too short");
+        return ask(`search:${s}`);
+      });
+      const results: () => Row[] = keyed
+        ? (() => {
+            const p = createProjection(() => src(), [] as Row[], { key: "id" });
+            return () => p;
+          })()
+        : src;
+      const show = createMemo(() => q() !== "");
+      const out = createMemo(() => {
+        if (!show()) return undefined;
+        return untrack(() =>
+          createErrorBoundary(
+            () =>
+              createLoadingBoundary(
+                () => mapArray(results, r => r.name),
+                () => "loading"
+              ),
+            e => `E:${(e() as Error).message}`
+          )
+        );
+      });
+      const resolve = (v: unknown): unknown =>
+        typeof v === "function" ? resolve(v()) : Array.isArray(v) ? v.join("") : v;
+      createRenderEffect(
+        () => resolve(out()),
+        v => void (screen = v)
+      );
+      return d;
+    });
+    await tick();
+    expect(screen).toBe(undefined);
+    setQ("a");
+    await tick();
+    expect(screen).toBe("E:too short");
+    setQ("ap");
+    await tick();
+    expect(screen).toBe("E:too short");
+    await land("search:ap", [
+      { id: 1, name: "apple" },
+      { id: 2, name: "apricot" }
+    ]);
+    expect(screen).toBe("appleapricot");
+    dispose();
+  });
diff --git a/packages/web/test/errored-born-staged-error-projection-4517.spec.tsx b/packages/web/test/errored-born-staged-error-projection-4517.spec.tsx
new file mode 100644
index 0000000000..ccd46a1784
--- /dev/null
+++ b/packages/web/test/errored-born-staged-error-projection-4517.spec.tsx
@@ -0,0 +1,79 @@
+/**
+ * @jsxImportSource @solidjs/web
+ * @vitest-environment jsdom
+ */
+import { expect, test } from "vitest";
+import { render } from "../src/index.js";
+import {
+  createMemo,
+  createProjection,
+  createSignal,
+  Errored,
+  flush,
+  For,
+  Loading,
+  Show
+} from "solid-js";
+
+type Row = { id: number; name: string };
+
+const tick = async () => {
+  for (let i = 0; i < 4; i++) {
+    await new Promise(r => setTimeout(r, 0));
+    flush();
+  }
+};
+
+test.each([true, false])(
+  "an answer replaces the error a boundary was born with (projection: %s)",
+  async projection => {
+    const asks: { key: string; res: (v: Row[]) => void }[] = [];
+    const ask = (key: string) => new Promise<Row[]>(res => asks.push({ key, res }));
+    const land = async (key: string, v: Row[]) => {
+      for (const a of asks.filter(a => a.key === key)) {
+        asks.splice(asks.indexOf(a), 1);
+        a.res(v);
+      }
+      await tick();
+    };
+    const container = document.createElement("div");
+    const [q, setQ] = createSignal("");
+    const dispose = render(() => {
+      const src = createMemo(() => {
+        const s = q();
+        if (s === "") return [] as Row[];
+        if (s.length < 2) throw new Error("too short");
+        return ask(`search:${s}`);
+      });
+      const results: () => Row[] = projection
+        ? (() => {
+            const p = createProjection(() => src(), [] as Row[], { key: "id" });
+            return () => p;
+          })()
+        : src;
+      return (
+        <Show when={q() !== ""}>
+          <Errored fallback={(e: () => any) => <p>{e().message}</p>}>
+            <Loading fallback={<p>loading</p>}>
+              <For each={results()}>{r => <i>{r.name}</i>}</For>
+            </Loading>
+          </Errored>
+        </Show>
+      );
+    }, container);
+    await tick();
+    expect(container.textContent).toBe("");
+    setQ("a");
+    await tick();
+    expect(container.textContent).toBe("too short");
+    setQ("ap");
+    await tick();
+    expect(container.textContent).toBe("too short");
+    await land("search:ap", [
+      { id: 1, name: "apple" },
+      { id: 2, name: "apricot" }
+    ]);
+    expect(container.textContent).toBe("appleapricot");
+    dispose();
+  }
+);